7.3.7
Causes of Speciation
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Allopatric Speciation
Allopatric speciation is a type of speciation where two (or more) populations are geographically isolated from each other.

Geographical isolation
- Populations of a species can split into two or more populations.
- This may be caused by environmental changes (e.g. emergence of a river) or random events (e.g. a storm causing a population of birds to be carried to another island).
- When two populations are physically separated, they are geographically isolated.

Selection pressures
- Populations in different environments will be exposed to different environmental conditions (e.g. climate, food availability).
- The different environments will place different selection pressures on the populations.
- The different selection pressures will drive natural selection in different directions.

Allele frequencies
- The different selection pressures on the populations cause the allele frequencies in the two populations to change.
- If the populations continue to be geographically isolated for a long time, eventually the allele frequencies will change so much that if the populations are brought back together they can no longer interbreed.

Speciation
- Species are defined as a group of actually or potentially interbreeding individuals.
- If two populations that have been geographically isolated can no longer interbreed when they are brought back together, they are considered to be two species.
- This is allopatric speciation.
Sympatric Speciation
Sympatric speciation is a type of speciation where two (or more) populations are not geographically isolated from each other.

Reproductive isolation
- In sympatric speciation, a population of species is reproductively isolated without geographical separation.
- Reproductive isolation may arise where a mutation emerges in a population that prevents some members of the population from interbreeding with others.

Reproductive isolation mutations
- Sympatric speciation could be caused by mutations that influence many different factors:
- Polyploidy - where a cell or organism has an extra set, or sets, of chromosomes. Polyploidy organisms are reproductively isolated from diploid organisms.
- Mutations may influence the flowering times or mating times of individuals.
- Mutations may alter the reproductive organs.
- Mutations may alter mating behaviours.

Speciation
- The biological definition of a species is a group of individuals that can actually or potentially interbreed.
- The emergence of a mutation that prevents two (or more) populations from interbreeding is called sympatric speciation.
Genetic Drift
Speciation can also be driven by genetic drift. Genetic drift is the random change in allele frequencies in a population.

Random changes in alleles
- By chance, some alleles are passed onto offspring and some are not.
- This causes random changes in the allele frequencies in the population.
- This is called genetic drift.

Speciation
- When two populations are reproductively isolated, they are exposed to different selection pressures. These selection pressures drive the evolution of the two populations in different directions. This is called disruptive selection.
- Genetic drift also contributes to the speciation by causing random changes in the allele frequencies of the two populations.
- Because the two populations can no longer interbreed, the gene pools will become increasingly different. This causes speciation.

Small populations
- Genetic drift tends to have a larger influence on evolutionary change in smaller populations.
- This is because the gene pool (the sum of all the genes of a population) is smaller.
- If the gene pool is smaller, any change in allele frequencies has a larger impact on the overall gene pool.
1Biological Molecules
1.1Monomers & Polymers
1.2Carbohydrates
1.3Lipids
1.4Proteins
1.4.1The Peptide Chain1.4.2Investigating Proteins1.4.3Primary & Secondary Protein Structure1.4.4Tertiary & Quaternary Protein Structure1.4.5Enzymes1.4.6Factors Affecting Enzyme Activity1.4.7Enzyme-Controlled Reactions1.4.8End of Topic Test - Lipids & Proteins1.4.9A-A* (AO3/4) - Enzymes1.4.10A-A* (AO3/4) - Proteins1.4.11Diagnostic Misconceptions - Enzyme Inhibitors
1.5Nucleic Acids
1.6ATP
1.7Water
1.8Inorganic Ions
2Cells
2.1Cell Structure
2.2Mitosis & Cancer
2.3Transport Across Cell Membrane
2.4Cell Recognition & the Immune System
2.4.1Immune System2.4.2Phagocytosis2.4.3T Lymphocytes2.4.4B Lymphocytes2.4.5Antibodies2.4.6Primary & Secondary Response2.4.7Vaccines2.4.8HIV2.4.9Ethical Issues2.4.10End of Topic Test - Immune System2.4.11Exam-Style Question - Immune System2.4.12A-A* (AO3/4) - Immune System2.4.13Diagnostic Misconceptions - Humoral vs Cellular
3Substance Exchange
3.1Surface Area to Volume Ratio
3.2Gas Exchange
3.3Digestion & Absorption
3.4Mass Transport
3.4.1Haemoglobin3.4.2Oxygen Transport3.4.3The Circulatory System3.4.4The Heart3.4.5Blood Vessels3.4.6Cardiovascular Disease3.4.7Heart Dissection3.4.8Xylem3.4.9Phloem3.4.10Investigating Plant Transport3.4.11End of Topic Test - Mass Transport3.4.12A-A* (AO3/4) - Mass Transport3.4.13Diagnostic Misconceptions - Concentration Gradient3.4.14Diagnostic Misconceptions - Cardiac Cycle3.4.15Diagnostic Misconceptions - Carrying Capacity3.4.16Diagnostic Misconceptions - Translocation
4Genetic Information & Variation
4.1DNA, Genes & Chromosomes
4.2DNA & Protein Synthesis
4.3Mutations & Meiosis
4.4Genetic Diversity & Adaptation
4.5Species & Taxonomy
4.6Biodiversity Within a Community
4.7Investigating Diversity
5Energy Transfers (A2 only)
5.1Photosynthesis
5.1.1Overview of Photosynthesis5.1.2Photoionisation of Chlorophyll5.1.3Production of ATP & Reduced NADP5.1.4Cyclic Photophosphorylation5.1.5Light-Independent Reaction5.1.6A-A* (AO3/4) - Photosynthesis Reactions5.1.7Limiting Factors5.1.8Photosynthesis Experiments5.1.9End of Topic Test - Photosynthesis5.1.10A-A* (AO3/4) - Photosynthesis
5.2Respiration
5.2.1Overview of Respiration5.2.2Anaerobic Respiration5.2.3A-A* (AO3/4) - Anaerobic Respiration5.2.4The Link Reaction5.2.5The Krebs Cycle5.2.6Oxidative Phosphorylation5.2.7Respiration Experiments5.2.8End of Topic Test - Respiration5.2.9A-A* (AO3/4) - Respiration5.2.10Diagnostic Misconceptions - Aerobic vs Anaerobic
5.3Energy & Ecosystems
6Responding to Change (A2 only)
6.1Nervous Communication
6.2Nervous Coordination
6.3Muscle Contraction
6.4Homeostasis
6.4.1Overview of Homeostasis6.4.2Blood Glucose Concentration6.4.3Controlling Blood Glucose Concentration6.4.4End of Topic Test - Blood Glucose6.4.5Primary & Secondary Messengers6.4.6Diabetes Mellitus6.4.7Measuring Glucose Concentration6.4.8Osmoregulation6.4.9Controlling Blood Water Potential6.4.10ADH6.4.11End of Topic Test - Diabetes & Osmoregulation6.4.12A-A* (AO3/4) - Homeostasis6.4.13Diagnostic Misconceptions - Effect of ADH
7Genetics & Ecosystems (A2 only)
7.1Genetics
7.2Populations
7.3Evolution
7.3.1Variation7.3.2Natural Selection & Evolution7.3.3End of Topic Test - Populations & Evolution7.3.4Types of Selection7.3.5Types of Selection Summary7.3.6Overview of Speciation7.3.7Causes of Speciation7.3.8Diversity7.3.9End of Topic Test - Selection & Speciation7.3.10A-A* (AO3/4) - Populations & Evolution7.3.11Diagnostic Misconceptions - Types of Speciation
8The Control of Gene Expression (A2 only)
8.1Mutation
8.2Gene Expression
8.2.1Stem Cells8.2.2Stem Cells in Disease8.2.3End of Topic Test - Mutation & Gene Epression8.2.4A-A* (AO3/4) - Mutation & Stem Cells8.2.5Regulating Transcription8.2.6Epigenetics8.2.7Epigenetics & Disease8.2.8Regulating Translation8.2.9Experimental Data8.2.10End of Topic Test - Transcription & Translation8.2.11Tumours8.2.12Correlations & Causes8.2.13Prevention & Treatment8.2.14End of Topic Test - Cancer8.2.15A-A* (AO3/4) - Gene Expression & Cancer
8.3Genome Projects
Jump to other topics
1Biological Molecules
1.1Monomers & Polymers
1.2Carbohydrates
1.3Lipids
1.4Proteins
1.4.1The Peptide Chain1.4.2Investigating Proteins1.4.3Primary & Secondary Protein Structure1.4.4Tertiary & Quaternary Protein Structure1.4.5Enzymes1.4.6Factors Affecting Enzyme Activity1.4.7Enzyme-Controlled Reactions1.4.8End of Topic Test - Lipids & Proteins1.4.9A-A* (AO3/4) - Enzymes1.4.10A-A* (AO3/4) - Proteins1.4.11Diagnostic Misconceptions - Enzyme Inhibitors
1.5Nucleic Acids
1.6ATP
1.7Water
1.8Inorganic Ions
2Cells
2.1Cell Structure
2.2Mitosis & Cancer
2.3Transport Across Cell Membrane
2.4Cell Recognition & the Immune System
2.4.1Immune System2.4.2Phagocytosis2.4.3T Lymphocytes2.4.4B Lymphocytes2.4.5Antibodies2.4.6Primary & Secondary Response2.4.7Vaccines2.4.8HIV2.4.9Ethical Issues2.4.10End of Topic Test - Immune System2.4.11Exam-Style Question - Immune System2.4.12A-A* (AO3/4) - Immune System2.4.13Diagnostic Misconceptions - Humoral vs Cellular
3Substance Exchange
3.1Surface Area to Volume Ratio
3.2Gas Exchange
3.3Digestion & Absorption
3.4Mass Transport
3.4.1Haemoglobin3.4.2Oxygen Transport3.4.3The Circulatory System3.4.4The Heart3.4.5Blood Vessels3.4.6Cardiovascular Disease3.4.7Heart Dissection3.4.8Xylem3.4.9Phloem3.4.10Investigating Plant Transport3.4.11End of Topic Test - Mass Transport3.4.12A-A* (AO3/4) - Mass Transport3.4.13Diagnostic Misconceptions - Concentration Gradient3.4.14Diagnostic Misconceptions - Cardiac Cycle3.4.15Diagnostic Misconceptions - Carrying Capacity3.4.16Diagnostic Misconceptions - Translocation
4Genetic Information & Variation
4.1DNA, Genes & Chromosomes
4.2DNA & Protein Synthesis
4.3Mutations & Meiosis
4.4Genetic Diversity & Adaptation
4.5Species & Taxonomy
4.6Biodiversity Within a Community
4.7Investigating Diversity
5Energy Transfers (A2 only)
5.1Photosynthesis
5.1.1Overview of Photosynthesis5.1.2Photoionisation of Chlorophyll5.1.3Production of ATP & Reduced NADP5.1.4Cyclic Photophosphorylation5.1.5Light-Independent Reaction5.1.6A-A* (AO3/4) - Photosynthesis Reactions5.1.7Limiting Factors5.1.8Photosynthesis Experiments5.1.9End of Topic Test - Photosynthesis5.1.10A-A* (AO3/4) - Photosynthesis
5.2Respiration
5.2.1Overview of Respiration5.2.2Anaerobic Respiration5.2.3A-A* (AO3/4) - Anaerobic Respiration5.2.4The Link Reaction5.2.5The Krebs Cycle5.2.6Oxidative Phosphorylation5.2.7Respiration Experiments5.2.8End of Topic Test - Respiration5.2.9A-A* (AO3/4) - Respiration5.2.10Diagnostic Misconceptions - Aerobic vs Anaerobic
5.3Energy & Ecosystems
6Responding to Change (A2 only)
6.1Nervous Communication
6.2Nervous Coordination
6.3Muscle Contraction
6.4Homeostasis
6.4.1Overview of Homeostasis6.4.2Blood Glucose Concentration6.4.3Controlling Blood Glucose Concentration6.4.4End of Topic Test - Blood Glucose6.4.5Primary & Secondary Messengers6.4.6Diabetes Mellitus6.4.7Measuring Glucose Concentration6.4.8Osmoregulation6.4.9Controlling Blood Water Potential6.4.10ADH6.4.11End of Topic Test - Diabetes & Osmoregulation6.4.12A-A* (AO3/4) - Homeostasis6.4.13Diagnostic Misconceptions - Effect of ADH
7Genetics & Ecosystems (A2 only)
7.1Genetics
7.2Populations
7.3Evolution
7.3.1Variation7.3.2Natural Selection & Evolution7.3.3End of Topic Test - Populations & Evolution7.3.4Types of Selection7.3.5Types of Selection Summary7.3.6Overview of Speciation7.3.7Causes of Speciation7.3.8Diversity7.3.9End of Topic Test - Selection & Speciation7.3.10A-A* (AO3/4) - Populations & Evolution7.3.11Diagnostic Misconceptions - Types of Speciation
8The Control of Gene Expression (A2 only)
8.1Mutation
8.2Gene Expression
8.2.1Stem Cells8.2.2Stem Cells in Disease8.2.3End of Topic Test - Mutation & Gene Epression8.2.4A-A* (AO3/4) - Mutation & Stem Cells8.2.5Regulating Transcription8.2.6Epigenetics8.2.7Epigenetics & Disease8.2.8Regulating Translation8.2.9Experimental Data8.2.10End of Topic Test - Transcription & Translation8.2.11Tumours8.2.12Correlations & Causes8.2.13Prevention & Treatment8.2.14End of Topic Test - Cancer8.2.15A-A* (AO3/4) - Gene Expression & Cancer
8.3Genome Projects
Practice questions on Causes of Speciation
Can you answer these? Test yourself with free interactive practice on Seneca — used by over 10 million students.
- 1
- 2Which of these is NOT essential for allopatric speciation?Multiple choice
- 3What is the definition of a species?Multiple choice
- 4
- 5Which of these does NOT cause sympatric speciation?Multiple choice
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