3.4.9

Phloem

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Structure

The phloem are plant vessels that are responsible for transporting sugars. The structure of the phloem is specialised for this role.

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Sieve tube elements

  • Sieve tube elements are the cells that make up the phloem vessel.
  • The vessel consists of many sieve tube elements that run up the stem on top of one another.
  • Sieve tube elements are living cells that contain a cytoplasm but no nucleus.
  • The walls of sieve tube elements are made of cellulose.
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Companion cells

  • Companion cells are connected to sieve tube elements via plasmodesmata.
    • Plasmodesmata allows the cytoplasm to be shared between companion cells and sieve tube elements.
  • Companion cells have a nucleus.
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Sieve plates

  • At either end of the sieve tube elements are sieve plates.
  • Sieve plates have large pores that allow sap to move through the sieve tube elements.
  • Sieve plates allow sugars to be transported through the phloem.

Translocation

Translocation is the process where sugars produced in photosynthesis are transported from the leaves to other parts of a plant. The steps involved are:

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1) Active transport of sucrose

  • Sucrose is produced during photosynthesis in the leaves.
    • The leaves are called the source.
  • Sucrose is actively transported into the companion cells in the phloem.
  • Energy is provided for active transport from mitochondria in the companion cells.
  • This process involves H+ ions being actively transported out of companion cells.
  • The H+ ions then diffuse back into the companion cells with sucrose. This is known as co-transport.
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2) Diffusion of sucrose

  • The loading of sucrose into the phloem causes the concentration of sucrose to increase.
  • Sucrose diffuses from the companion cells into the sieve tube elements.
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3) Osmosis

  • The increase in the concentration of sucrose in the phloem causes water potential in the phloem to decrease.
  • There is a water potential gradient between the outside of the phloem and the inside.
  • Water diffuses into the phloem by osmosis.
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4) Unloading of sucrose

  • At the bottom of the phloem, sucrose concentration is low because it is being used up in the cells.
    • The cells at the bottom of the phloem are called the sink.
  • Sucrose diffuses out of the phloem and into the sink cells.
  • This lowers the water potential of the sink cells.
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5) Osmosis

  • Water diffuses down its water potential gradient out of the phloem by osmosis.
  • The diffusion of water into the phloem at the source and out of the phloem at the sink creates a hydrostatic pressure gradient.
  • The hydrostatic pressure gradient allows mass transport of sucrose from source to sink.
    • This is called mass flow.

Jump to other topics

1Biological Molecules

2Cells

3Substance Exchange

4Genetic Information & Variation

5Energy Transfers (A2 only)

6Responding to Change (A2 only)

7Genetics & Ecosystems (A2 only)

8The Control of Gene Expression (A2 only)

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