3.4.8

Xylem

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Structure

The xylem are plant vessels that are responsible for transporting water and mineral ions. The structure of the xylem is specialised for this role.

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Vessel elements

  • The xylem vessels are long, tubes of cells that run up the stem of plants.
  • The cells of the xylem are called vessel elements.
  • The vessel elements in the xylem are dead.
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End walls

  • The vessel elements are stacked on top one another.
  • There are no cell walls at the ends of each vessel element.
  • This creates a continuous tube for water to flow through.
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Lignin

  • The walls of the xylem are lined with a waterproof polymer called lignin.
  • Lignin reinforces the walls of the vessel elements to provide structural support.

Cohesion-Tension Theory

Water and inorganic ions travel up the xylem through cohesion and tension. The steps involved in this process are:

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1) Transpiration

  • Some of the water in the leaves is used in photosynthesis.
  • Most of the water in the leaves evaporates in a process called transpiration.
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2) Tension

  • The loss of water from the leaves creates tension in the xylem.
  • Tension is the formation of hydrogen bonds between water molecules and the sides of the xylem vessel elements.
  • Water in the xylem is pulled upwards by this tension towards the leaves.
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3) Cohesion

  • Individual water molecules also form hydrogen bonds with each other. This process is called cohesion.
  • When water molecules are pulled up the xylem, other molecules of water are also pulled upwards due to cohesion.
  • The combination of cohesion and tension together continuously pull water upwards to replace water that has been lost in the leaves by transpiration.
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4) Diffusion in the roots

  • When water is pulled up the stem, the water potential at the bottom of the plant decreases.
  • Water diffuses into the roots via osmosis down its water potential gradient.

Investigating Transpiration Rate

The rate of transpiration can be influenced by a number of factors. An experiment can be set up using a potometer to estimate transpiration rate. The steps involved are:

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1) Assemble the potometer

  • A potometer is a piece of equipment that is used to estimate the volume of water taken up by a plant in a given time.
  • The potometer is filled with water and a cutting of a shoot is placed inside.
  • The shoot must be cut and placed into the potometer while underwater to ensure no air enters the xylem.
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2) Form the air bubble

  • Remove the end of the capillary tube from the water beaker.
  • Wait for a bubble of air to form in the capillary tube.
  • Place the capillary tube back into the water.
  • The air bubble is used to record the volume of water used by the shoot.
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3) Record bubble movement

  • Mark the starting position of the air bubble.
  • Use a stopwatch to record the distance moved by the air bubble in a given time period.
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4) Calculate transpiration rate

  • Calculate the rate of movement of the bubble per hour.
  • The rate of bubble movement is equal to the transpiration rate.
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5) Change the variable

  • The experiment can be repeated by changing a different variable each time (e.g. temperature or light).
  • It is important to only change one variable at one time and to keep all other conditions constant.
  • Changing one variable allows the effects of an environmental factor on transpiration rate to be compared.

Jump to other topics

1Biological Molecules

2Cells

3Substance Exchange

4Genetic Information & Variation

5Energy Transfers (A2 only)

6Responding to Change (A2 only)

7Genetics & Ecosystems (A2 only)

8The Control of Gene Expression (A2 only)

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