14.5.1

Stomata & Guard Cells

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Stomata & Guard Cells

Plants regulate gaseous exchange through their stomata. Stomata are apertures found predominantly on the underside of leaves. Each stomatal aperture is formed by two guard cells.

Daily rhythm

Daily rhythm

  • Stomata regulate gaseous exchange and water loss through transpiration.
  • Stomata tend to be open during the day to allow gaseous exchange for photosynthesis. Plants need CO2 for photosynthesis.
  • But, having stomata open means water is lost through transpiration.
  • So, stomata tend to close at night to prevent excessive water loss.
    • There is also no sunlight at night, so the plant has no requirement for carbon dioxide for respiration.
Guard cells

Guard cells

  • Two guard cells form a stomatal aperture.
  • Guard cells have a thick cell wall in their middle portion. So, when guard cells are full of water (turgid) they bend away from the thick middle portion.
    • Each turgid guard cell forms a "C" shape, so the stomatal aperture is open.
  • Following water loss, guard cells become flaccid. Flaccid guard cells are linear and lie side-by-side.
  • This closes the guard cell, preventing further water loss.
Regulating water

Regulating water

  • Potassium (K+) ions are used to regulate guard cell opening.
  • Turgidity is caused by an increase in K+ ion concentration within guard cells.
  • A higher K+ ion concentration lowers the water potential of the guard cell. Water then follows by osmosis.
  • This makes guard cells turgid and opens the stomatal aperture.
  • Stomata close following excessive water loss. This is usually due to a lower rate of photosynthesis following loss of sunlight.
Water stress

Water stress

  • Water stress is communicated in plants by the hormone abscisic acid (ABA).
  • The roots of plants produce ABA in response to stress or low water potentials.
  • ABA travels in water to guard cells. Once in guard cells, ABA triggers the release of Ca2+ ions.
  • Ca2+ ions act as a second messenger. It binds to and activates proteins.
    • Ca2+ ions initiate a signalling cascade that results in the stomatal aperture closing.
Jump to other topics
1

Cell Structure

2

Biological Molecules

3

Enzymes

4

Cell Membranes & Transport

5

The Mitotic Cell Cycle

6

Nucleic Acids & Protein Synthesis

7

Transport in Plants

8

Transport in Mammals

9

Gas Exchange

10

Infectious Diseases

11

Immunity

12

Energy & Respiration (A2 Only)

13

Photosynthesis (A2 Only)

14

Homeostasis (A2 Only)

15

Control & Coordination (A2 Only)

16

Inherited Change (A2 Only)

17

Selection & Evolution (A2 Only)

18

Classification & Conservation (A2 Only)

19

Genetic Technology (A2 Only)

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